Main Ref. | Ribbink, A.J., B.A. Marsh, A.C. Marsh, A.C. Ribbink and B.J. Sharp, 1983 |
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Remarks | Inhabits the intermediate zone. Also found in sediment covered slabs and small medium rocks. Territorial males, usually centered around a hole among the rocks in which the resident spawns or takes refuge, ritualized intraspecific fights common. Females, juveniles and non territorial adult males occur in large schools, frequently numbering hundreds of individuals as well as singly or in small groups. Non territorial fish feed in the water column, territorial males may also do when plankton is available. Sand, rock or sediment scrapers. |
Marine - Neritic | Marine - Oceanic | Brackishwater | Freshwater | |
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Marine zones / Brackish and freshwater bodies |
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Substrate | |
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Substrate Ref. | |
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Special habitats Ref. |
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Parasitism |
Feeding type | plants/detritus+animals (troph. 2.2-2.79) |
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Feeding type Ref. | Ribbink, A.J., B.A. Marsh, A.C. Marsh, A.C. Ribbink and B.J. Sharp, 1983 |
Feeding habit | grazing on aquatic plants |
Feeding habit Ref. | Ribbink, A.J., B.A. Marsh, A.C. Marsh, A.C. Ribbink and B.J. Sharp, 1983 |
Estimation method | Original sample | Unfished population | Remark | ||
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Troph | s.e. | Troph | s.e. | ||
From diet composition | 2.26 | 0.10 | 2.52 | 0.09 | Troph of juv./adults from 4 studies. |
From individual food items | |||||
Ref. | Ribbink, A.J., B.A. Marsh, A.C. Marsh, A.C. Ribbink and B.J. Sharp, 1983 |