Phalloceros anisophallos Lucinda, 2008
Roque toothcarp
photo by Lucinda, P.H.F.

Family:  Poeciliidae (Poeciliids), subfamily: Poeciliinae
Max. size:  2.82 cm SL (male/unsexed); 4.51 cm SL (female)
Environment:  benthopelagic; freshwater
Distribution:  South America: Brazil in the drainages of rio Parati, rio Barra Grande, rio São Roque, rio Taquari and rio Itinguçu (small coastal drainages of Rio de Janeiro).
Diagnosis:  Dorsal soft rays (total): 7-8; Anal soft rays: 9-11; Vertebrae: 32-32. Autapomorphically diagnosed by having hook on left half of gonopodial appendix near its tip, located approximately from distal quarter to the distal third of length of gonopodial appendix, and spot on median region of flanks elongate, forming a vertical bar reaching dorsal and ventral profiles. Differs from its congeners but Phalloceros megapolos, Phalloceros spiloura, Phalloceros malabarbai, Phalloceros alessandrae, Phalloceros buckupi, Phalloceros uai, Phalloceros lucenorum, Phalloceros pellos, and Phalloceros reisi by the female urogenital papilla curved to the right, located laterally (vs. slightly left turned and with a lateral ramus or straight located along midventral line); and border of the anal aperture in contact with the first anal-fin ray or very close to it (vs. separated from first anal-fin ray by the female urogenital papilla). Can be differentiated from its congeners except Phalloceros spiloura, Phalloceros uai, Phalloceros lucenorum, Phalloceros pellos, and Phalloceros reisi by the absence of the right hook and presence of the left one (vs. absence or presence of left and right hooks), and gonopodial appendix asymmetrical; its halves different from each other; right half wider than left one (vs. gonopodial appendix symmetrical; its halves similar to each other). Can be separated from Phalloceros spiloura by the absence of a caudal peduncle spot (vs. presence of caudal peduncle spot); from Phalloceros uai by the presence of a wide and square-shaped lateral spot in large specimens (vs. absence of such spot); from Phalloceros lucenorum by rectangular tip of hook on left half of gonopodial appendix (vs. rounded tip); from Phalloceros lucenorum, Phalloceros pellos, and Phalloceros reisi by the lateral spot in adult females very narrow, forming a vertically elongated bar covering up to the length corresponding to two scales length in horizontal direction and 2-4 scales in vertical direction (vs. lateral spot in adult females absent, small, or elliptical to roundish, but never forming a vertical bar) (Ref. 76852). Description: Pectoral fin with 5-7 branched rays; longitudinal series of scales 28-29; transverse series of scales 6-7 (Ref. 76852).
Biology: 
IUCN Red List Status: Least Concern (LC); Date assessed: 07 November 2018 Ref. (130435)
Threat to humans:  harmless


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