Metamorphosed females distinguished by the following characteristics: globular escal bulb, surrounded distally and laterally by a blunt ridge in small specimens; absence of escal filaments or other appendages; barbel divided at base into 4 primary branches of nearly equal length, 40-86% SL, each with varying number of secondary and tertiary branches, more or less prolonged into filaments, each with distal photophore; concentration of subdermal melanophores on caudal peduncle; males with pointed sphenotic spines (Ref. 86949).