Can be diagnosed by the following characters: mouth subterminal; oral valve reaching the anterior edge of vomer; two posterior nasal pores, posterior one smaller; suborbital pores seven (6 + 1); lateral line double with ventral and medio-lateral branches; supraoccipital dividing the posterior end of frontals; basal pectoral plate with one foramen between scapular strut and r1; r2 and r3 notched; single row of palatine teeth; dorsal fin rays 109-112; anal fin rays 89-93; pectoral fin rays 17; vertebrae asymmetrical, 27 + 89-92 = 116-119; gill slit extending ventrally to just lower end of pectoral fin base; two pyloric caeca well developed; pseudobranch filaments 3, elongated. Scales, ventral fins and vomerine teeth present (Ref. 89950).