You can sponsor this page

Raja clavata Linnaeus, 1758

Thornback ray
Add your observation in Fish Watcher
Native range | All suitable habitat | Point map | Year 2050
This map was computer-generated and has not yet been reviewed.
Raja clavata   AquaMaps   Data sources: GBIF OBIS
Upload your photos and videos
Pictures | Videos | Google image
Image of Raja clavata (Thornback ray)
Raja clavata
Picture by Stergiou, K.I.

Classification / Names Nomi Comuni | Sinonimi | Catalog of Fishes(Genere, Specie) | ITIS | CoL | WoRMS | Cloffa

Elasmobranchi (squali e razze) (sharks and rays) > Rajiformes (Skates and rays) > Rajidae (Skates)
Etymology: Raja: Latin, raja, -ae = a sting ray (Raja sp.) (Ref. 45335).
More on author: Linnaeus.

Environment: milieu / climate zone / depth range / distribution range Ecologia

marino demersale; distribuzione batimetrica 5 - 1020 m (Ref. 114953). Subtropical; 70°N - 38°S, 25°W - 50°E

Distribuzione Stati | Aree FAO | Ecosystems | Presenze | Point map | Introduzioni | Faunafri

Eastern Atlantic and Southwest Indian Ocean: Iceland to Madagascar, including the Mediterranean Sea.

Length at first maturity / Size / Peso / Age

Maturity: Lm 76.6, range 47 - 87.5 cm
Max length : 105 cm TL maschio/sesso non determinato; (Ref. 53748); 139.0 cm TL (female); common length : 85.0 cm TL maschio/sesso non determinato; (Ref. 9987); peso massimo pubblicato: 18.0 kg (Ref. 35388); Età massima riportata: 15 anni (Ref. 41305)

Short description Chiavi di identificazione | Morfologia | Morfometria

Spine dorsali (totale) : 0; Spine anali: 0. Sub-rhomboid in shape with pointed wing-tips; disc-width 1,25 to 1,36 times in its length, its length 1,70 to 1,83 times in total length. Short rostrum, rounded at his extremity; pectoral fins with clear angles on lateral side; triangular pelvic fins (Ref. 39246). Dorsally prickly; large females also prickly throughout their ventral surface; juveniles and large males prickly along the margins of their discs and the underside of their snout. 30-50 thorns form a median row from the nape to the first dorsal fin; additional large 'buckler' thorns with swollen bases scattered on upper surface of disc in adults (Ref. 3167). Coloration is highly variable, with the dorsal surface having all shades of brown with dark and light spots and blotches; ventral surface white (Ref. 3167, 58137).

Biologia     Glossario (es. epibenthic)

Inhabits shelf and upper slope waters. Reported depth ranges varies, from 10-300 m (Ref. 4426, 82319, 88187), and from 300-577 m in the eastern Ionian Sea (Ref. 56504); most common in coastal waters between 10-60 m depth (Ref. 58137). Tolerates low salinities (Ref. 88171). Found on mud, sand and gravel bottoms, rarely on rougher bottoms (Ref. 6808, 58137). Nocturnal species. Feeds on all kinds of bottom animals, preferably crustaceans (Ref. 3167) and fishes (Ref. 114953). Undertakes migrations with mean distances of 54-117 km per month; shows a clear annual migration cycle (Ref. 89017, 89018), moves from deeper offshore waters (10-30 m) in autumn and winter to shallower areas (<10 m) in spring (Ref. 82319). Young are non-migratory, inhabiting inshore nursery grounds (Ref. 89019, 89020, 89022); in the Bay of Douarnenez (France) they are found to remain in shallow waters for at least 2 years (Ref. 89023). Maturity age 7-8 years (Ref. 114953). Oviparous. Young may tend to follow large objects, such as their mother (Ref. 205). Detects weak electric fields generated by other organisms (prey detection and predator avoidance) but may also generate its own weak electric fields (Ref. 10311, 88171). Utilized fresh and frozen (Ref. 9987).

Life cycle and mating behavior Maturities | Riproduzione | Spawnings | Egg(s) | Fecundities | Larve

Oviparous. Distinct pairing with embrace. Polyandrous species (Ref. 89024). Paired eggs are laid and deposited on shallow sand, mud, pebble or gravel bottoms (Ref. 205, 89025). Up to 170 egg cases can be laid by a single female in a year (Ref. 3167), but average fecundity is much lower (around 48-74 eggs) (Ref. 3603, 31302). In northwestern Europe, egg cases are laid during spring (Ref. 3167) and in the Mediterranean during winter and spring (Ref. 3167). Egg cases are oblong capsules with stiff pointed horns at the corners, each containing one embryo (Ref. 205). Capsules are 5.0-9.0 cm long without the horns (Ref. 41250, 88187) and 3.4-6.8 cm wide (Ref. 41250). Egg cases are anchored with an adhesive film (Ref. 82399). Embryos feed solely on yolk (Ref. 50449). Egg cases hatch after about 4-5 months and pups are about 11-13 cm TL (Ref. 88864). Mating season from February to September, peaking in June (Ref. 74501). Adults observed to form same-sex aggregations during the mating season with females moving to shallower inshore waters approximately a month before the males (Ref 3603, 58137, 74501). Mating does not occur in the Baltic Sea (Ref. 82311).

Main reference Upload your references | Bibliografia | Coordinatore : McEachran, John | Collaboratori

McEachran, J.D. and K.A. Dunn, 1998. Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia 1998(2):271-290. (Ref. 27314)

IUCN Red List Status (Ref. 130435)

  Near Threatened (NT) ; Date assessed: 03 October 2005

CITES

Not Evaluated

CMS (Ref. 116361)

Not Evaluated

Threat to humans

  Harmless





Human uses

Pesca: commerciale; Pesce da pesca sportiva: si
FAO - pesca: landings; Publication: search | FishSource | Sea Around Us

Informazioni ulteriori

Trophic ecology
Prede
Diet compositions
Food consumptions
Food rations
Predatori
Ecology
Ecologia
Life cycle
Riproduzione
Maturities
Fecundities
Spawnings
Spawning aggregations
Egg(s)
Egg developments
Larve
Dinamica popolazioni larvali
Distribution
Stati
Aree FAO
Ecosystems
Presenze
Introduzioni
BRUVS - Videos
Anatomy
Gill areas
Cervelli
Otoliths
Physiology
Body compositions
Nutrients
Oxygen consumptions
Swimming type
Swimming speeds
Visual pigment(s)
Suoni dei Pesci
Diseases / Parasites
Toxicities (LC50s)
Genetics
Genetica
Electrophoreses
Heritabilities
Human related
Aquaculture systems
Profili di acquacoltura
Varietà
Ciguatera cases
Stamps, coins, misc.
Outreach
Collaboratori
References
Bibliografia

Strumenti

Special reports

Download XML

Fonti Internet

AFORO (otoliths) | Aquatic Commons | BHL | Cloffa | BOLDSystems | Websites from users | Check FishWatcher | CISTI | Catalog of Fishes: Genere, Specie | DiscoverLife | DORIS | ECOTOX | FAO - pesca: landings; Publication: search | Faunafri | Fishipedia | Fishtrace | GenBank: genome, nucleotide | GloBI | Google Books | Google Scholar | Google | IGFA World Record | MitoFish | Database Nazionali | Otolith Atlas of Taiwan Fishes | Acquari pubblici | PubMed | Reef Life Survey | Socotra Atlas | Tree of Life | Wikipedia: Go, ricerca | World Records Freshwater Fishing | Zoobank | Zoological Record

Estimates based on models

Preferred temperature (Ref. 123201): 7.1 - 15.8, mean 10.7 °C (based on 474 cells).
Phylogenetic diversity index (Ref. 82804):  PD50 = 0.5000   [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00200 (0.00175 - 0.00228), b=3.26 (3.23 - 3.29), in cm total length, based on LWR estimates for this species (Ref. 93245).
Trophic level (Ref. 69278):  3.8   ±0.2 se; based on diet studies.
Generation time: 7.8 (5.4 - 10.4) years. Estimated as median ln(3)/K based on 16 growth studies.
Resilienza (Ref. 120179):  Basso, tempo minimo di raddoppiamento della popolazione 4.5 - 14 anni (K=0.09-0.14; tm=10; tmax=23; Fec=150).
Prior r = 0.18, 95% CL = 0.12 - 0.26, Based on 4 full stock assessments.
Fishing Vulnerability (Ref. 59153):  High vulnerability (60 of 100).
Climate Vulnerability (Ref. 125649):  High vulnerability (60 of 100).
Price category (Ref. 80766):   Medium.
Nutrients (Ref. 124155):  Calcium = 9.22 [1.18, 176.74] mg/100g; Iron = 0.628 [0.060, 7.651] mg/100g; Protein = 17.2 [14.4, 19.5] %; Omega3 = 0.571 [0.265, 1.295] g/100g; Selenium = 39.7 [7.5, 183.1] μg/100g; VitaminA = 5.22 [0.40, 59.91] μg/100g; Zinc = 0.47 [0.03, 5.30] mg/100g (wet weight); based on nutrient studies.